Abundance: 0.05 to 5.15% (median 0.20%)

Latitudinal range: 40.9° to 50.6°

Habitats: (7), temperate broadleaf/mixed forest (1)

Found in 8 samples

France: Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene)

Greece: Lake Prespa National Park

Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 17) (Early Pleistocene), Zabia Cave (Layer 15) (Early Pleistocene)

See also Vipera, Vipera ammodytes, Vipera aspis, Vipera lebetina, Vipera natiensis, Vipera palaestinae, Vipera sp., Vipera ursinii